|DATA ON NAME:|
|Name||Flabellophora kinabaluensis Corner|
|Protologue||Nova Hedwigia, Beih. 86: 28|
|Information on type||Type: on fallen trunks in montane forest, Malaysia, Sabah|
|Current name||Flabellophora kinabaluensis Corner|
|Classification||Polyporaceae, Polyporales, Agaricomycetes, Basidiomycota|
Flabellophora kinabaluensis sp. nov. Figure 2, Plate 10 Borneo, Mt. Kinabalu, 1600-1800 m alt., frequens. Typus RSNB 5614 in herb. Corner.
Merismatoid -14 cm wide. Pilei -5 cm in radius, flabelliform, oblique, imbricating, matt, becoming very unevenly rugoso-tuberculate, not zoned or slightly near the margin, white then isabelline-drab, becoming slightly fawn-drab or subfusous towards the white subacute margin. Trunk more or less central, short, stout, -3 cm thick, dividing into shortly stipitate and more or less pleuropodal pilei, concolorous. Tubes -2.5 mm long, white; pores 150-250 Ám wide, angular, entire, dissepiments 100-180 Ám thick, white. Flesh 4-10 mm thick at the base of the pilei, coriaceous, hard in the trunk, somewhat watery zoned or marbled, white, not discolouring on bruising. Smell rather fragrant. On fallen trunks in montane forest, 1600-1800 m alt. Borneo, Mt. Kinabalu, frequent.
Spores 4.5-5 x 2.5-3 Ám (?larger), white, smooth, ellipsoid, not dextrinoid. Basidia 18-22 x 5.5-6 Ám clavate; sterigmata 4, 4-5 Ám long. Cystidia and hyphal pegs none. Hymenium not thickening. Hyphae dimitic with intercalary skeletal cells, the tissue becoming very densely interwoven, not dextrinoid, not encrusted; generative hyphae 3-9 Ám wide, clamped, becoming somewhat thick-walled, with frequent H-connections, often branched from the clamp, developing in the older tissue (especially above the tubes and below the surface of the pileus) intricate knots of narrow, rather thick-walled, branches 1.5-5 Ám wide, simulating binding hyphae but with clamps; skeletals as intercalary cells -1200 Ám long, 4-9 Ám wide, radiating and interwoven, with 0-2 branches near the distal end, walls -2.5 Ám thick, lumen often linear, the middle wall swelling in potash but not diffluent; in the dissepiments truly dimitic with unbranched, unlimited skeletals 2.5-3.5 Ám wide, the generative hyphae 1.5-5 Ám wide. Surface of pileus with a close pile of cylindric hyphal ends -120 x 4-6 Ám, becoming thick-walled towards their bases, weathering away and drying up but the stalks of these hyphal ends becoming subagglutinated without forming a distinct crust.
Collections: Bembangan R., 1600 m alt., 3 March 1964, RSNB 5614; Mesilau; 1800 m alt., 7 Feb. 1964, RSNB 5268 (young).
In young specimens (RSNB 5268) the pileus was violaceous towards the margin. The colour disappears in mature specimens and the tissue becomes so dense that its construction can hardly be made out. The merismatoid habit represents a condensed state of that in F. superposita, but F. kinabaluensis can also be regarded as a merismatoid Tyromyces. Indeed, I have several very similar species from this montane district of Kinabalu which have the sessile dimidiate fruitbodies of Tyromyces and can be separated genrically from Flabellophora only by the absence of a stem. F. kinabaluensis seems to represent the progenitor of one group of Tyromyces.
Flabellophora kinabaluensis sp. nov. Figure 2, Plate 10 Merismatoidea -14 cm lata. Pilei -5 cm in radio, flabelliformes subascendentes imbricati, sublaeves dein basim versus rugoso-tuberculati, pallide albi dein isabellini subcervini vel subfusci; marginibus albis. Truncus -3 cm crassus, plus minus mesopodalis, in pileis pleuropodalibus breviter stipitatis divisus, concolor. Tubi -2.5 mm longi, albi; poris 150-250 Ám latis, angulatis, integris, albis. Care 4-10 mm crassa ad basim pilei, coriacea, in trunco dura, fractu haud discolorans, plus minus aquoso-zonata, alba. Odor subfragrans. Sporae 4.5-5 x 2.5-3 Ám, albae laeves ellipsoideae, haud dextrinoideae. Cystidia nulla. Hyphae dimiticae, nec dextrinoideae nec incrustatae; generatrices 3-9 Ám latae, fibulatae, e fibula saepe ramosae; skeletales intercalares -1200 Ám longae, tunica -2.5 Ám incrassata; carne vetusta ramis generatricibus ut ligativis, saepe in nodulis contortis; in dissepimentis hyphae 1.5-5 Ám latae, dense intertextae, skeletales 2.5-3.5 Ám latae et haud ramosae. Ad truncos emortuos in silva montana. Borneo, Mt. Kinabalu, 1600-1800 m alt., frequens. Typus RSNB 5614 in herb. Corner.
Flabellophora kinabaluensis Corner, Beih. Nova Hedwig. 86:28 (1987). Holotype: MALAYSIA, Borneo, Mt. Kinabalu, Bemgangan R, 3 Mar. 1964, leg. E. J. H. Corner (E). Accepted as Grifola kinabaluensis (Corner) Hattori comb. nov. (Basionym indicated above.) This species is placed temporarily in Grifola Gray, because of the merismatoid basidiocarps and dimitic hyphal system with clamp connections, as its real taxonomic position is unclear.
Basidiocarps merismatoid, pilei flabelliform, imbricated. Pileus surface velutinous to glabrous, azonate, warted near the base, irregularly wrinkled near the margin, grayish orange (5 A-B 4-5) to dark brown (7-8 D-F 8). Pores angular, 3-4/mm, dissepiments thin, eroded. Context leathery to corky, somewhat horny, brittle near the margin, with agglutinated crust, orange white (5 A 2). Tubes brittle, concolorous with the context.
Trama hyphae dimitic: generative hyphae with clamp connections, thin-walled, hyaline, 2-5 Ám wide; skeletal hyphae thick-walled with a distinct lumen, hyaline, 1.5-3 Ám wide. Context hyphae subdimitic: generative hyphae with clamp connections, thin- to slightly thickwalled, 2.5-7 Ám wide; skeletal hyphae thick-walled with a distinct lumen, occasionally flattened, hyaline, IKI-, 2-10 Ám wide, somewhat fragile in Melzer's reagent, swelled in 5% KOH solution. Basidiospores not seen.
|Reference to description||Hattori, T. 2000, Mycoscience 41(4): 346|
|Comments or additions please to: J. Stalpers||Date: May 26, 2013|